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Identification of Bean goose subspecies



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  • 江田真毅,嶋田哲郎,溝田智俊,小池裕子.2011.脱落羽毛の遺伝的解析からみた日本で越冬するヒシクイ Anser fabalis の亜種区分.日本鳥学会誌.60(1), 100-104.

  • Fox, A.D. & Leafloor, J.O. (eds.) 2018. A Global Audit of the Status and Trends of Arctic and Northern Hemisphere Goose Populations (Component 2: Population accounts). Conservation of Arctic Flora and Fauna International Secretariat: Akureyri, Iceland. ISBN 978-9935-431-74-5

  • Heinicke, T. 2011. Status of the Bean Goose Anser fabalis wintering in central Asia. Wildfowl. 59: 77-99.

  • 池内俊雄.2017.ヒシクイ・オオヒシクイQ&A(四訂版).雁の里親友の会.pp40

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  • 呉地正行,横田義雄,大津真理子.1983.ヒシクイAnser fabalis serrirostrisとオオヒシクイA. f. middendorfiの野外識別についての考察.Tori,32:95-108

  • Miyabayashi, Y. & Mundkur, T. 1999. Atlas of Key Sites for Anatidae in the East Asian Flyway. Wetlands International - Japan, Tokyo, and Wetlands International - Asia Pacific, Kuala Lumpur, 148pp. [on-line]

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  • Ottenburghs, J., Honka, J., Müskens, G.J.D.M. et al. Recent introgression between Taiga Bean Goose and Tundra Bean Goose results in a largely homogeneous landscape of genetic differentiation. Heredity 125, 73–84 (2020).

  • Reeber, S. 2015. Waterfowl of North America, Europe & Asia: An identification guide. Princeton University Press. 656pp

  • Ruokonen M., Litvin K. & Aarvak T. 2008. Taxonomy of the bean goose – pink-footed goose complex. Molecular Phylogenetics and Evolution 48, 554–562.

  • Ruokonen, M. and Aarvak, K. 2011. Typology revisited: historical taxa of the bean goose – pink-footed goose complex. Ardea 99(1), 103-112

  • Sangster, G., Oreel, G.J., 1996. Progress in taxonomy of taiga and tundra bean geese. Dutch Bird. 18, 310–316

  • Wetlands International (2019). "Waterbird Population Estimates" . Retrieved from on Thursday 28 Feb 2019






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